SCIENCE15 - Color Perception

Pat suggested that I write a blog about color perception after she read an article that proposed that women may have an advantage over men.  So, I looked into the subject, and found that color perception is a very complicated subject.  But it was also a very fascinating subject, so I decided to go ahead, trying not to make it too technical.   

 

After a short introduction, I will talk about the nature of color, how we see color, why individuals may not see the same color, and end with the importance of color to our lives today.

My principal sources include: “Color Vision,” Wikipedia.com; “Primary Colors - What Are the Primary Colors in Color Theory?” artincontext.org; “Why We Don’t See the Same Colors,” psychologytoday.com; “The Science of How We See Color,” datacolor.com; “Colour,” Britannica.com; “Color is in the eye, and brain, of the beholder,” knowablemagazine.org; “Color Blindness:  The Most Common, Uncommon Eye Condition,” downtownvisionnv.com; plus, numerous other online sources.

 

Introduction 

Color vision is a feature of visual perception that allows us perceive a multitude of different colors.

In humans and other primates, color vision may have evolved under selective pressure for a variety of visual tasks including foraging for nutritious young leaves, ripe fruit, and flowers, as well as detecting predator camouflage and emotional states in other primates.

Today, color helps us remember objects, influences our purchases, and sparks our emotions. 

Objects do not possess color.  They reflect wavelengths of light that are seen as color by the human brain.  When white light strikes a white object, it appears white to us because it absorbs no color and reflects all color equally.  When it strikes a colored object, this color light is reflected back to our eyes.  A black object absorbs all colors equally and reflects none, so it looks black to us. 

A person can see in dim light without being able to distinguish colors.  Only when more light is present do colors appear.  Light of some critical intensity, therefore, is necessary for color perception.  Finally, the way the brain responds to visual stimuli must also be considered.  Even under identical conditions, the same object may appear red to one observer and orange to another.  Clearly, the perception of color depends on vision, light, and individual interpretation.

 

The Nature of Color

Aristotle viewed color to be the product of a mixture of white and black, and this was the prevailing belief until 1666, when Isaac Newton’s prism experiments provided the scientific basis for the understanding of color.  Newton showed that a glass prism could break up white light (like sunlight) into a range of colors, which he called the spectrum, and that the recombination of these spectral colors re-created the white light.  Although he recognized that the spectrum was continuous, Newton used the seven color names violet, indigo, blue, green, yellow, orange, and red for segments of the spectrum (some say by analogy with the seven notes of the musical scale).

In 1666, Isaac Newton showed that a prism could break up white light into a range of colors.


 

The Visible Light Spectrum.  Today, we know that color is associated specifically with electromagnetic radiation of a certain range of wavelengths visible to the human eye.   This range of wavelengths is known as the visible spectrum - i.e., light. 

Electromagnetic radiation is energy that travels at the speed of light and spreads out as it goes.  From short wavelengths to long wavelengths, the types of electronic radiation that make up the electromagnetic spectrum include:  cosmic rays, X-rays, ultraviolet light, visible light, infrared light, microwaves, radar, radio, and the broadcast band consisting of TV and AM and FM radio.

The figure below shows the electromagnetic spectrum and the narrow range of visible light, extending left to right, from violet light to red light. 

The visible light spectrum is a narrow range of the electromagnetic spectrum.

 

The visible light spectrum for humans ranges from about 380 to 740 nanometers (a nanometer is one billionth of a meter).  Newton’s spectral colors, with cyan (blue-green) replacing indigo, can be found in this range.

These spectral colors do not refer to a single wavelength, but rather to a set of wavelengths: violet, 380 - 450 nm; blue, 450 - 485 nm; cyan, 485 - 500 nm; green, 500 - 565 nm; yellow, 565 - 590 nm; orange, 590 - 625 nm; red, 625 - 740 nm. 

Wavelengths just shorter and longer than this visible range are ultraviolet light and infrared light, respectively.  Humans cannot generally see these wavelengths, but some animals can. 

Spectral Colors. A spectral color can be precisely specified by its hue, saturation, and brightness - three attributes sufficient to distinguish it from all other possible perceived colors.

Hue is the attribute of colors that permits them to be classed as red, yellow, green, blue, or an intermediate between any contiguous pair of these colors.  Hue is the wavelength within the visible-light spectrum at which the reflected energy output from a viewed object is greatest.

Saturation refers to relative purity.  When a pure, vivid, strong shade of red is mixed with a variable amount of white, weaker or paler reds are produced, each having the same hue but a different saturation.  As saturation increases, colors appear sharper or purer.  As saturation decreases, colors appear more washed-out or faded. 

Finally, light of any given combination of hue and saturation can have a variable brightness (also called intensity), which depends on the total amount of light energy present.  The intensity of the color grows with increased brightness, but the color itself does not change.

Note:  Many photo processing computer programs have adjustable settings for these parameters.

Non-Spectral Colors.  There are a variety of colors in addition to spectral colors.  These include grayscale colors, shades of colors obtained by mixing grayscale colors with spectral colors, violet-red colors, and metallic colors.

Grayscale colors include white, gray, and black.

Shades include colors such as pink or Navy. Pink is obtained from mixing red and white.  Navy is obtained from mixing blue and black.

Violet-red colors include hues and shades of magenta.  The light spectrum is a line on which violet is one end and the other is red, and yet we see hues of purple that connect those two colors.

A metallic color is a color that appears to be that of a polished metal.  The visual sensation usually associated with metals is its metallic shine. 

Now that I’ve talked about the nature of color, I want to talk about how we see color.

 

How We See Color

We see colors thanks to photoreceptor cells in the retinas of our eyes that transmit signals to our brains. 

Rods and Cones.  Light enters the human eye, where the lens focuses the light on the retina, the layer of nerve cells in the back of the eye.  The retina is covered by millions of light-sensitive cells, some shaped like rods and some like cones.  These receptors process the light into nerve impulses and pass them along to the cortex of the brain via the optic nerve.

We see colors thanks to photoreceptor rods and cones in our eyes that transmit signals to our brains.

 

Rods work at very low levels of light.  We use these for night vision because only a few bits of light (photons) can activate a rod.  Rods don't help with color vision, which is why at night, we see everything in a gray scale. 

Cones require a lot more light, and they are used to see color.  We have three types of cones: blue (short wavelengths), green (medium wavelengths), and red (long wavelengths), that contain different forms of opsin - a pigment protein - that has different sensitivities to light. 

As shown in the graph below, each cone type is able to detect a range of colors (wavelengths).  Even though each cone type is most sensitive to a specific color of light (where the line peaks), they also can detect other colors (shown by the stretch of each curve). 

It is the overlap of the cone types, and how the brain integrates and interprets the signals sent from them, that allows us to see millions of different colors. 

Each of the three human cone types is able to detect a range of colors.

 

Note:  This is the so-called RGB color model, associated with the wavelengths of visible light.  The primary colors are red, green, and blue, from which all other spectral colors can be obtained with appropriate mixing. (See the figure at the beginning of this article.)  The traditional color theory we all learned in school, tells us that the primary colors are red, yellow, and blue.  That color theory only involves the use of pigments in paint, and does not take into consideration the way light blends color. 

 Human peripheral vision is less sharp and colorful than front-on vision.  That’s because of the rods and cones.  Rods are most highly concentrated around the edge of the retina. There are over 120 million of them in each eye.  Rods transmit mostly black and white information to the brain.  As rods are more sensitive to dim light than cones, we lose most color vision in dusky light and our peripheral vision is less colorful.  It is the rods that help our eyes adjust when we enter a darkened room.

Cones are concentrated in the middle of the retina, with fewer on the periphery.  Six million cones in each eye transmit the higher levels of light intensity that create the sensation of color and visual sharpness. 

Dimension.  There is another property of color perception called “dimension,” which has to do with the number of different cone types. 

Most humans have three cone types, which absorb maximally in different regions of the wavelength spectrum.  So, most humans are trichromats.  However, eight percent of males (and an insignificant number of females) have only two cone types. They are dichromats (color-deficient).

The complete list of possible dimensions, based on the number of different cone types present, is:

Monochromacy - lack of any color perception.  Color-blind.  Many mammals, such as cetaceans (whales, dolphins, porpoises), the owl monkey, and the Australian sea lion are monochromats.

Dichromacy - most mammals and a quarter of color-blind humans have this color vision deficiency.

Trichromacy - most humans.  All perceptible colors can be formed with different combinations of the primary colors:  red, green, and blue.

Tetrachromacy - most birds, reptiles, and fish.

Pentachromacy - rare in vertebrates.  Some birds, notably pigeons; some lampreys (jawless fish).

Some insects, especially bees, can see ultraviolet colors invisible to the human eye. 

Color Blindness.  Color blindness doesn’t have anything to do with how sharp our vision is or how much light we see, but it does mean the cone cells process colors differently.  It affects 1 in 12 men throughout their life, and is much rarer in women, where only 1 in 200 will have some kind of color deficiency.  It’s often inherited genetically through the mother but has been known to develop over time with age or diseases like diabetes and multiple sclerosis.  Color blindness can be separated into three different categories: red-green color blindness, blue-yellow color blindness, and the much rarer complete color blindness.

 

Why Individuals Don’t See the Same Color

A lot of factors feed into how people perceive and talk about color, from the biology of our eyes to how our brains process that information, to the words our languages use to talk about color categories.  There’s plenty of room for differences, all along the way.

For example, most people have three types of light receptor cones in the eye that are optimized to detect different wavelengths or colors of light.  But sometimes, a genetic variation can cause one type of cone to be different, or absent altogether, leading to altered color vision.  Some people are color-blind.  Others may have color superpowers (see below).

Our sex can also play a role in how we perceive color, as well as our age, and even the color of our irises.  Our perception can change depending on where we live, when we were born, and what season it is.

It would be rather surprising if there were no variation in how we experience colors.  The number of cones in the human retina is not the same for everyone.  Sometimes individuals have many cones, and sometimes they are barely present.  This difference has been observed in so-called normal individuals who react in the same way to color stimuli.

The fact that the number of cones in our eyes varies considerably suggests that the brain must be able to automatically adjust the input from the retina.  So, individual variations in color perception may not purely be a matter of the nature and number of the cones in the retina.  It can also be a result of the fact that people with different numbers of cones calibrate the input from the retina in different ways.

Recent studies indicate significant variance in a gene located on the X chromosome which codes for a protein that detects light in the long-wavelength (red/orange) regions of the color spectrum.  Since women have two copies of the X chromosome, it is possible for them to have two different versions of this gene, and hence it is possible for them to have a more fine-grained ability to discriminate light in the long-wavelength regions of the color spectrum.  Women are thus potentially able to perceive a broader spectrum of colors (color vision superpower) in the long-wavelength regions than men.

 

Importance of Color to Our Lives Today

Color symbolism serves important roles in art, religion, politics, and ceremonials, as well as in everyday life.

The most important aspect of color in daily life is probably the one that is least defined and most variable.  It involves aesthetic and psychological responses to color and influences art, fashion, commerce, and even physical and emotional sensations.  One example of the link between color and emotion is the common perception that red, orange, yellow, and brown hues are “warm,” while the blues, greens, and grays are “cold.”  The red, orange, and yellow hues are said to induce excitement, cheerfulness, stimulation, and aggression; the blues and greens induce security, calm, and peace; and the browns, grays, and blacks induce sadness, depression, and melancholy. 

Many psychologists believe that analyzing an individual’s uses of and responses to color can reveal information about the individual’s physiological and psychological condition. It has even been suggested that specific colors can have a therapeutic effect on physical and mental disabilities.

Although medical benefits are still in question, color has been shown to cause definite physical and emotional reactions in humans and in some animals.  Rooms and objects that are white or in light shades of “cool” colors may appear to be larger than those that are in intense dark or “warm” colors; black or very dark colors have a slimming, or shrinking, effect, as is well known to designers and decorators.  A “cool” room decorated in a pale blue requires a higher thermostat setting than a “warm” room painted a pale orange to achieve the same sensation of warmth.  People who view a display of unusual colors produced by special illumination may experience headaches and nervous disorders; tasty wholesome food served under such conditions appears repulsive and may even induce illness.  Some colors induce a feeling of pleasure in the observer.

The effect of combinations of colors on an observer depends not only on the individual effects of the colors but also on the harmony of the colors combined and the composition of the pattern.  Artists and designers have been studying the effects of colors for centuries and have developed a multitude of theories on the uses of color.  The number and variety of these theories demonstrate that no universally accepted rules apply; the perception of color depends on individual experience.

Colors even play a vital role in our safety.  Like the yellow school bus. Why is it important that we see it, even in our periphery?  For safety, of course.  Many colors are used to depict important safety messages without words.  Red stop signs and green traffic lights are universal.



 

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